Flavitrack Analysis of the Structure and Function of West Nile Non-structural Proteins

Petr Danecek and Catherine Schein



The Flavitrack database was developed to relate the sequences of the Flaviviridae to the phenotype of the viruses, to aid in design of vaccines and specific antivirals. Statistical methods were developed to compare the genomes of the Flaviviruses, which are evolutionarily closely related organisms with high subtype variability. Due to the variability within each virus group, methods that compare randomly selected representatives from each of the virus groups can give misleading results. Here, statistically rigorous methods were used to analyze all available sequences for each group separately. The resulting consensus sequences were then used to calculate motifs based on conserved physicochemical properties. In the application illustrated here, the tools were used to compare 919 sequences of NS2a, one of the viral proteins that block the host interferon response to infection. The analysis indicated that residues, which if altered in Kunjin (West Nile) sequence, reduced the ability of the virus to grow and prevented it from blocking interferon production, are all in the conserved N-terminus. One possible model, based on a nuclear receptor protein structure, placed three of these residues (I59, V61, M103) close to one another in a negatively charged pocket on the protein surface.

Index Terms Consensus sequence, Flavitrack, NS2a, PCPMer, West Nile virus.